Colorado PROFILES, The Colorado Clinical and Translational Sciences Institute (CCTSI)
Keywords
Last Name
Institution

MIchael James Franklin

TitleInstructor/Fellow
InstitutionUniversity of Colorado Denver - Anschutz Medical Campus
DepartmentSOM-PSYCH

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    Publications listed below are automatically derived from MEDLINE/PubMed and other sources, which might result in incorrect or missing publications. Faculty can login to make corrections and additions.
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    1. Williamson KS, Franklin MJ. Role of starvation survival response mechanisms on ribosome integrity, antibiotic tolerances, and virulence of Pseudomonas aeruginosa biofilms. NPJ Biofilms Microbiomes. 2026 Jul 08; 12(1). PMID: 42414298.
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    2. Welland MJ, Ahlquist KD, De Fazio P, Austin-Tse C, Pais L, Wedd L, Bryen S, Rius R, Franklin M, Morrison C, Hall G, Gauthier L, Bloemendal A, Francis DI, Mallett AJ, Mallawaarachchi A, Lockhart PJ, Leventer R, Scheffer IE, Howell KB, Kassahn KS, Scott HS, McGaughran J, Christodoulou J, Thorburn DR, Thompson BA, Patel CV, Smith G, O'Donnell-Luria A, Sadedin S, Rehm HL, Lunke S, Wander J, Samocha KE, Simons C, MacArthur DG, Stark Z. Automated reanalysis of genomic data for rare disease diagnostics at scale. Nat Med. 2026 Aug; 32(8):2991-2999. PMID: 42343115.
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    3. Williamson KS, Franklin MJ. An optimized mung bean seedling model for characterizing virulence of Pseudomonas aeruginosa biofilm infections. Biofilm. 2026 Jun; 11:100363. PMID: 42318084.
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    4. Salpadoru T, Khanam S, Borin VA, Achour MA, Oh D, Kanik M, Gallage PC, Khanov A, Hull M, Pitre SP, Agarwal PK, Franklin MJ, Patrauchan MA. The PA2803-encoded PcrP exhibits a novel non-catalytic function and contributes to polymyxin B resistance in P. aeruginosa. J Bacteriol. 2025 11 20; 207(11):e0018925. PMID: 41104944.
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    5. Welland MJ, Ahlquist KD, De Fazio P, Austin-Tse C, Pais L, Wedd L, Bryen S, Rius R, Franklin M, Morrison C, Hall G, Gauthier L, Bloemendal A, Francis DI, Mallett AJ, Mallawaarachchi A, Lockhart PJ, Leventer R, Scheffer IE, Howell KB, Kassahn KS, Scott HS, McGaughran J, Christodoulou J, Thorburn DR, Thompson BA, Patel CV, Smith G, O'Donnell-Luria A, Sadedin S, Rehm HL, Lunke S, Wander J, Samocha KE, Simons C, MacArthur DG, Stark Z. Scalable automated reanalysis of genomic data in research and clinical rare disease cohorts. medRxiv. 2025 May 21. PMID: 40661289.
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    6. Salpadoru T, Khanam S, Borin VA, Achour MA, Oh D, Kanik M, Gallage PC, Khanov A, Hull M, Pitre S, Agarwal PK, Franklin MJ, Patrauchan MA. A novel non-catalytic function of PA2803-encoded PcrP contributes to polymyxin B resistance in P. aeruginosa and redefines the functional role of the PA2803 subfamily. bioRxiv. 2025 May 13. PMID: 40462998.
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    7. Braciszewski JM, Colby SM, Franklin MJ, Stout RL, Vose-O'Neal A, Bock BC. Perspectives on a Technology-Based Smoking Cessation Intervention Among Youth Exiting Foster Care. Child Youth Serv Rev. 2025 Jun; 173. PMID: 40463359.
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    8. Tanudisastro HA, Cuomo ASE, Weisburd B, Welland M, Spenceley E, Franklin M, Xue A, Bowen B, Wing K, Tang O, Gray M, Reis ALM, Margoliash J, Kurtas NE, Pullin JM, Lee AS, Brand H, Harper M, Bobowik K, Silk M, Marshall J, Bakiris V, Madala BS, Uren C, Bartie C, Senabouth A, Dashnow H, Fearnley L, Martin-Trujillo A, Dolzhenko E, Qiao Z, Grieve SM, Nguyen T, Ben-David E, Chen L, Farh KK, Talkowski M, Alexander SI, Siggs OM, Gruenschloss L, Nicholas HR, Piscionere J, Simons C, Wallace C, Gymrek M, Deveson IW, Hewitt AW, Figtree GA, de Lange KM, Powell JE, MacArthur DG. Polymorphic tandem repeats influence cell type-specific gene expression across the human immune landscape. bioRxiv. 2025 Apr 09. PMID: 40291654.
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    9. Smith HJ, Franklin MJ. Use of epifluorescence widefield deconvolution microscopy for imaging and three-dimensional rendering of Pseudomonas aeruginosa biofilms and extracellular matrix materials. Methods Microbiol. 2023; 53:309-324. PMID: 38974073.
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    10. Williamson KS, Dlakic M, Akiyama T, Franklin MJ. The Pseudomonas aeruginosa RpoH (s32) Regulon and Its Role in Essential Cellular Functions, Starvation Survival, and Antibiotic Tolerance. Int J Mol Sci. 2023 Jan 12; 24(2). PMID: 36675051.
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    11. Braciszewski JM, Colby SM, Franklin MJ, Stout RL, Vose-O'Neal A, Bock BC. Technology-Based Smoking Cessation for Youth Exiting Foster Care: A Pilot Randomized Trial. J Public Child Welf. 2024; 18(1):61-79. PMID: 38144460.
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    12. Stewart PS, Williamson KS, Boegli L, Hamerly T, White B, Scott L, Hu X, Mumey BM, Franklin MJ, Bothner B, Vital-Lopez FG, Wallqvist A, James GA. Search for a Shared Genetic or Biochemical Basis for Biofilm Tolerance to Antibiotics across Bacterial Species. Antimicrob Agents Chemother. 2022 04 19; 66(4):e0002122. PMID: 35266829.
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    13. Potter DA, Herrera-Ponzanelli CA, Hinojosa D, Castillo R, Hernandez-Cruz I, Arrieta VA, Franklin MJ, Yee D. Recent advances in neoadjuvant therapy for breast cancer. Fac Rev. 2021; 10:2. PMID: 33659921.
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    14. Theng S, Williamson KS, Franklin MJ. Role of Hibernation Promoting Factor in Ribosomal Protein Stability during Pseudomonas aeruginosa Dormancy. Int J Mol Sci. 2020 Dec 14; 21(24). PMID: 33327444.
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    15. Franklin MJ, Sandvik E, Yanardag S, Williamson KS. Functional Characterization of the Pseudomonas aeruginosa Ribosome Hibernation-Promoting Factor. J Bacteriol. 2020 09 08; 202(19). PMID: 32900865.
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    16. Buechler CR, Ukani A, Elsharawi R, Gable J, Petersen A, Franklin M, Chung R, Bell J, Manly A, Hefzi N, Carpenter D, Bryce R. Barriers, beliefs, and practices regarding hygiene and vaccination among the homeless during a hepatitis A outbreak in Detroit, MI. Heliyon. 2020 Mar; 6(3):e03474. PMID: 32258449.
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    17. King MM, Kayastha BB, Franklin MJ, Patrauchan MA. Calcium Regulation of Bacterial Virulence. Adv Exp Med Biol. 2020; 1131:827-855. PMID: 31646536.
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    18. Stewart PS, White B, Boegli L, Hamerly T, Williamson KS, Franklin MJ, Bothner B, James GA, Fisher S, Vital-Lopez FG, Wallqvist A. Conceptual Model of Biofilm Antibiotic Tolerance That Integrates Phenomena of Diffusion, Metabolism, Gene Expression, and Physiology. J Bacteriol. 2019 11 15; 201(22). PMID: 31501280.
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    19. Pratt SL, Zath GK, Akiyama T, Williamson KS, Franklin MJ, Chang CB. DropSOAC: Stabilizing Microfluidic Drops for Time-Lapse Quantification of Single-Cell Bacterial Physiology. Front Microbiol. 2019; 10:2112. PMID: 31608020.
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    20. Old JM, Lin SH, Franklin MJM. Mapping out bare-nosed wombat (Vombatus ursinus) burrows with the use of a drone. BMC Ecol. 2019 09 18; 19(1):39. PMID: 31533684.
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    21. Hamner S, Brown BL, Hasan NA, Franklin MJ, Doyle J, Eggers MJ, Colwell RR, Ford TE. Metagenomic Profiling of Microbial Pathogens in the Little Bighorn River, Montana. Int J Environ Res Public Health. 2019 03 27; 16(7). PMID: 30934749.
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    22. Akiyama T, Williamson KS, Franklin MJ. Expression and regulation of the Pseudomonas aeruginosa hibernation promoting factor. Mol Microbiol. 2018 10; 110(2):161-175. PMID: 29885070.
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    23. Akiyama T, Williamson KS, Schaefer R, Pratt S, Chang CB, Franklin MJ. Resuscitation of Pseudomonas aeruginosa from dormancy requires hibernation promoting factor (PA4463) for ribosome preservation. Proc Natl Acad Sci U S A. 2017 03 21; 114(12):3204-3209. PMID: 28270601.
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    24. Jones BC, Wood JG, Chang C, Tam AD, Franklin MJ, Siegel ER, Helfand SL. A somatic piRNA pathway in the Drosophila fat body ensures metabolic homeostasis and normal lifespan. Nat Commun. 2016 12 21; 7:13856. PMID: 28000665.
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    25. James GA, Ge Zhao A, Usui M, Underwood RA, Nguyen H, Beyenal H, deLancey Pulcini E, Agostinho Hunt A, Bernstein HC, Fleckman P, Olerud J, Williamson KS, Franklin MJ, Stewart PS. Microsensor and transcriptomic signatures of oxygen depletion in biofilms associated with chronic wounds. Wound Repair Regen. 2016 03; 24(2):373-83. PMID: 26748963.
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    26. Smith HJ, Foreman CM, Akiyama T, Franklin MJ, Devitt NP, Ramaraj T. Genome Sequence of Janthinobacterium sp. CG23_2, a Violacein-Producing Isolate from an Antarctic Supraglacial Stream. Genome Announc. 2016 Jan 28; 4(1). PMID: 26823573.
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    27. Guragain M, King MM, Williamson KS, Pérez-Osorio AC, Akiyama T, Khanam S, Patrauchan MA, Franklin MJ. The Pseudomonas aeruginosa PAO1 Two-Component Regulator CarSR Regulates Calcium Homeostasis and Calcium-Induced Virulence Factor Production through Its Regulatory Targets CarO and CarP. J Bacteriol. 2016 Jan 11; 198(6):951-63. PMID: 26755627.
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    28. Franklin MJ, Chang C, Akiyama T, Bothner B. New Technologies for Studying Biofilms. Microbiol Spectr. 2015 Aug; 3(4). PMID: 26350329.
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    29. Thumma SC, Jacobson BA, Patel MR, Konicek BW, Franklin MJ, Jay-Dixon J, Sadiq A, De A, Graff JR, Kratzke RA. Antisense oligonucleotide targeting eukaryotic translation initiation factor 4E reduces growth and enhances chemosensitivity of non-small-cell lung cancer cells. Cancer Gene Ther. 2015 Aug; 22(8):396-401. PMID: 26227824.
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    30. Stewart PS, Franklin MJ, Williamson KS, Folsom JP, Boegli L, James GA. Contribution of stress responses to antibiotic tolerance in Pseudomonas aeruginosa biofilms. Antimicrob Agents Chemother. 2015 Jul; 59(7):3838-47. PMID: 25870065.
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    31. Chimileski S, Franklin MJ, Papke RT. Biofilms formed by the archaeon Haloferax volcanii exhibit cellular differentiation and social motility, and facilitate horizontal gene transfer. BMC Biol. 2014 Aug 14; 12:65. PMID: 25124934.
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    32. Sarkisova SA, Lotlikar SR, Guragain M, Kubat R, Cloud J, Franklin MJ, Patrauchan MA. A Pseudomonas aeruginosa EF-hand protein, EfhP (PA4107), modulates stress responses and virulence at high calcium concentration. PLoS One. 2014; 9(2):e98985. PMID: 24918783.
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    33. Ding F, Gil MP, Franklin M, Ferreira J, Tatar M, Helfand SL, Neretti N. Transcriptional response to dietary restriction in Drosophila melanogaster. J Insect Physiol. 2014 Oct; 69:101-6. PMID: 24819200.
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    34. Smith H, Akiyama T, Foreman C, Franklin M, Woyke T, Teshima H, Davenport K, Daligault H, Erkkila T, Goodwin L, Gu W, Xu Y, Chain P. Draft Genome Sequence and Description of Janthinobacterium sp. Strain CG3, a Psychrotolerant Antarctic Supraglacial Stream Bacterium. Genome Announc. 2013 Nov 21; 1(6). PMID: 24265494.
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    35. Hamner S, McInnerney K, Williamson K, Franklin MJ, Ford TE. Bile salts affect expression of Escherichia coli O157:H7 genes for virulence and iron acquisition, and promote growth under iron limiting conditions. PLoS One. 2013; 8(9):e74647. PMID: 24058617.
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    36. Lewis TA, Glassing A, Harper J, Franklin MJ. Role for ferredoxin:NAD(P)H oxidoreductase (FprA) in sulfate assimilation and siderophore biosynthesis in Pseudomonads. J Bacteriol. 2013 Sep; 195(17):3876-87. PMID: 23794620.
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    37. Franklin MJ. Recognizing the research achievements of new investigators. Transl Res. 2013 Aug; 162(2):130-1. PMID: 23769765.
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    38. Fabich HT, Vogt SJ, Sherick ML, Seymour JD, Brown JR, Franklin MJ, Codd SL. Microbial and algal alginate gelation characterized by magnetic resonance. J Biotechnol. 2012 Oct 31; 161(3):320-7. PMID: 22728394.
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    39. Williamson KS, Richards LA, Perez-Osorio AC, Pitts B, McInnerney K, Stewart PS, Franklin MJ. Heterogeneity in Pseudomonas aeruginosa biofilms includes expression of ribosome hibernation factors in the antibiotic-tolerant subpopulation and hypoxia-induced stress response in the metabolically active population. J Bacteriol. 2012 Apr; 194(8):2062-73. PMID: 22343293.
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    40. Zhang H, Li W, Franklin MJ, Dudek AZ. Polymorphisms in DNA repair gene XRCC1 and skin cancer risk: a meta-analysis. Anticancer Res. 2011 Nov; 31(11):3945-52. PMID: 22110224.
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    41. Warren AE, Boulianne-Larsen CM, Chandler CB, Chiotti K, Kroll E, Miller SR, Taddei F, Sermet-Gaudelus I, Ferroni A, McInnerney K, Franklin MJ, Rosenzweig F. Genotypic and phenotypic variation in Pseudomonas aeruginosa reveals signatures of secondary infection and mutator activity in certain cystic fibrosis patients with chronic lung infections. Infect Immun. 2011 Dec; 79(12):4802-18. PMID: 21930755.
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    42. Franklin MJ, Nivens DE, Weadge JT, Howell PL. Biosynthesis of the Pseudomonas aeruginosa Extracellular Polysaccharides, Alginate, Pel, and Psl. Front Microbiol. 2011; 2:167. PMID: 21991261.
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    43. Raza A, Franklin MJ, Dudek AZ. Pericytes and vessel maturation during tumor angiogenesis and metastasis. Am J Hematol. 2010 Aug; 85(8):593-8. PMID: 20540157.
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    44. Obulareddy SJ, Xin J, Truskinovsky AM, Anderson JK, Franklin MJ, Dudek AZ. Metanephric adenoma of the kidney: an unusual diagnostic challenge. Rare Tumors. 2010 Jun 30; 2(2):e38. PMID: 21139840.
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    45. Antony GK, Bertino E, Franklin M, Otterson GA, Dudek AZ. Small cell lung cancer in never smokers: report of two cases. J Thorac Oncol. 2010 May; 5(5):747-8. PMID: 20421768.
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    46. Pérez-Osorio AC, Williamson KS, Franklin MJ. Heterogeneous rpoS and rhlR mRNA levels and 16S rRNA/rDNA (rRNA gene) ratios within Pseudomonas aeruginosa biofilms, sampled by laser capture microdissection. J Bacteriol. 2010 Jun; 192(12):2991-3000. PMID: 20348255.
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    47. Dudek AZ, Murthaiah PK, Franklin M, Truskinovsky AM. Metastatic adamantinoma responds to treatment with receptor tyrosine kinase inhibitor. Acta Oncol. 2010; 49(1):101-4. PMID: 19412810.
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    48. Spector E, Franklin MJ, Truskinovsky AM, Dudek AZ. Sorafenib induces partial response in metastatic medullary thyroid carcinoma. Acta Oncol. 2010; 49(1):104-6. PMID: 19711206.
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    49. Dai T, Tegos GP, Lu Z, Huang L, Zhiyentayev T, Franklin MJ, Baer DG, Hamblin MR. Photodynamic therapy for Acinetobacter baumannii burn infections in mice. Antimicrob Agents Chemother. 2009 Sep; 53(9):3929-34. PMID: 19564369.
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    50. Kim J, Hahn JS, Franklin MJ, Stewart PS, Yoon J. Tolerance of dormant and active cells in Pseudomonas aeruginosa PA01 biofilm to antimicrobial agents. J Antimicrob Chemother. 2009 Jan; 63(1):129-35. PMID: 19001451.
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    51. Pérez-Osorio AC, Franklin MJ. qRT-PCR of Microbial Biofilms. CSH Protoc. 2008 Oct 01; 2008:pdb.prot5066. PMID: 21356697.
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    52. Pérez-Osorio AC, Franklin MJ. Isolation of RNA and DNA from biofilm samples obtained by laser capture microdissection microscopy. CSH Protoc. 2008 Oct 01; 2008:pdb.prot5065. PMID: 21356696.
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    53. Lenz AP, Williamson KS, Pitts B, Stewart PS, Franklin MJ. Localized gene expression in Pseudomonas aeruginosa biofilms. Appl Environ Microbiol. 2008 Jul; 74(14):4463-71. PMID: 18487401.
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    54. Stewart PS, Franklin MJ. Physiological heterogeneity in biofilms. Nat Rev Microbiol. 2008 Mar; 6(3):199-210. PMID: 18264116.
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    55. Nguyen S, Franklin M, Dudek AZ. Skin ulcers in Klippel-Trenaunay syndrome respond to sunitinib. Transl Res. 2008 Apr; 151(4):194-6. PMID: 18355766.
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    56. Patrauchan MA, Sarkisova SA, Franklin MJ. Strain-specific proteome responses of Pseudomonas aeruginosa to biofilm-associated growth and to calcium. Microbiology (Reading). 2007 Nov; 153(Pt 11):3838-3851. PMID: 17975093.
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    57. Hammerschmidt DE, Franklin M. The limits and power of peer review. Minn Med. 2006 Jun; 89(6):43-5. PMID: 16846184.
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    58. Hammerschmidt DE, Franklin MJ. The edible wolf peach. J Lab Clin Med. 2005 Oct; 146(4):251-2. PMID: 16194687.
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    59. Patrauchan MA, Sarkisova S, Sauer K, Franklin MJ. Calcium influences cellular and extracellular product formation during biofilm-associated growth of a marine Pseudoalteromonas sp. Microbiology (Reading). 2005 Sep; 151(Pt 9):2885-2897. PMID: 16151201.
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    60. Douthit SA, Dlakic M, Ohman DE, Franklin MJ. Epimerase active domain of Pseudomonas aeruginosa AlgG, a protein that contains a right-handed beta-helix. J Bacteriol. 2005 Jul; 187(13):4573-83. PMID: 15968068.
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    61. Sarkisova S, Patrauchan MA, Berglund D, Nivens DE, Franklin MJ. Calcium-induced virulence factors associated with the extracellular matrix of mucoid Pseudomonas aeruginosa biofilms. J Bacteriol. 2005 Jul; 187(13):4327-37. PMID: 15968041.
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    62. Hammerschmidt DE, Franklin M. Secrecy in medical journals. Minn Med. 2005 Mar; 88(3):34-5. PMID: 15852594.
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    63. Skoff RP, Bessert DA, Cerghet M, Franklin MJ, Rout UK, Nave KA, Carlock L, Ghandour MS, Armant DR. The myelin proteolipid protein gene modulates apoptosis in neural and non-neural tissues. Cell Death Differ. 2004 Dec; 11(12):1247-57. PMID: 15375385.
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    64. Werner E, Roe F, Bugnicourt A, Franklin MJ, Heydorn A, Molin S, Pitts B, Stewart PS. Stratified growth in Pseudomonas aeruginosa biofilms. Appl Environ Microbiol. 2004 Oct; 70(10):6188-96. PMID: 15466566.
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    65. Franklin MJ, Brusilow WS, Woodbury DJ. Determination of proton flux and conductance at pH 6.8 through single FO sectors from Escherichia coli. Biophys J. 2004 Nov; 87(5):3594-9. PMID: 15339819.
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    66. Franklin MJ, Douthit SA, McClure MA. Evidence that the algI/algJ gene cassette, required for O acetylation of Pseudomonas aeruginosa alginate, evolved by lateral gene transfer. J Bacteriol. 2004 Jul; 186(14):4759-73. PMID: 15231808.
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    67. Jesaitis AJ, Franklin MJ, Berglund D, Sasaki M, Lord CI, Bleazard JB, Duffy JE, Beyenal H, Lewandowski Z. Compromised host defense on Pseudomonas aeruginosa biofilms: characterization of neutrophil and biofilm interactions. J Immunol. 2003 Oct 15; 171(8):4329-39. PMID: 14530358.
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    68. Jain S, Franklin MJ, Ertesvåg H, Valla S, Ohman DE. The dual roles of AlgG in C-5-epimerization and secretion of alginate polymers in Pseudomonas aeruginosa. Mol Microbiol. 2003 Feb; 47(4):1123-33. PMID: 12581364.
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    69. Walters MC, Roe F, Bugnicourt A, Franklin MJ, Stewart PS. Contributions of antibiotic penetration, oxygen limitation, and low metabolic activity to tolerance of Pseudomonas aeruginosa biofilms to ciprofloxacin and tobramycin. Antimicrob Agents Chemother. 2003 Jan; 47(1):317-23. PMID: 12499208.
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    70. Franklin MJ, Ohman DE. Mutant analysis and cellular localization of the AlgI, AlgJ, and AlgF proteins required for O acetylation of alginate in Pseudomonas aeruginosa. J Bacteriol. 2002 Jun; 184(11):3000-7. PMID: 12003941.
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    71. Morea A, Mathee K, Franklin MJ, Giacomini A, O'Regan M, Ohman DE. Characterization of algG encoding C5-epimerase in the alginate biosynthetic gene cluster of Pseudomonas fluorescens. Gene. 2001 Oct 31; 278(1-2):107-14. PMID: 11707327.
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    72. Xu KD, Franklin MJ, Park CH, McFeters GA, Stewart PS. Gene expression and protein levels of the stationary phase sigma factor, RpoS, in continuously-fed Pseudomonas aeruginosa biofilms. FEMS Microbiol Lett. 2001 May 15; 199(1):67-71. PMID: 11356569.
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    73. Nivens DE, Ohman DE, Williams J, Franklin MJ. Role of alginate and its O acetylation in formation of Pseudomonas aeruginosa microcolonies and biofilms. J Bacteriol. 2001 Feb; 183(3):1047-57. PMID: 11208804.
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    74. Anderl JN, Franklin MJ, Stewart PS. Role of antibiotic penetration limitation in Klebsiella pneumoniae biofilm resistance to ampicillin and ciprofloxacin. Antimicrob Agents Chemother. 2000 Jul; 44(7):1818-24. PMID: 10858336.
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    75. Keswani J, Orkand S, Premachandran U, Mandelco L, Franklin MJ, Whitman WB. Phylogeny and taxonomy of mesophilic Methanococcus spp. and comparison of rRNA, DNA hybridization, and phenotypic methods. Int J Syst Bacteriol. 1996 Jul; 46(3):727-35. PMID: 8782682.
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    76. Franklin MJ, Ohman DE. Identification of algI and algJ in the Pseudomonas aeruginosa alginate biosynthetic gene cluster which are required for alginate O acetylation. J Bacteriol. 1996 Apr; 178(8):2186-95. PMID: 8636017.
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    77. Franklin MJ, Chitnis CE, Gacesa P, Sonesson A, White DC, Ohman DE. Pseudomonas aeruginosa AlgG is a polymer level alginate C5-mannuronan epimerase. J Bacteriol. 1994 Apr; 176(7):1821-30. PMID: 8144447.
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    78. Franklin MJ, Ohman DE. Identification of algF in the alginate biosynthetic gene cluster of Pseudomonas aeruginosa which is required for alginate acetylation. J Bacteriol. 1993 Aug; 175(16):5057-65. PMID: 8394313.
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    79. Franklin MJ, Wiebe WJ, Whitman WB. Populations of methanogenic bacteria in a georgia salt marsh. Appl Environ Microbiol. 1988 May; 54(5):1151-7. PMID: 16347628.
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