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Richard Kellogg Olson

TitleSr Research Associate
InstitutionUniversity of Colorado Boulder
DepartmentInstitute for Behav Genetics
Phone303/492-8865

    Collapse Research 
    Collapse research activities and funding
    R01HD038526     (OLSON, RICHARD K)Mar 15, 2000 - May 31, 2013
    NIH
    Longitudinal Twin Study--Early Reading Development
    Role: Principal Investigator
    P50HD027802     (WILLCUTT, ERIK G)Sep 30, 1990 - Jul 31, 2022
    NIH
    Differential Diagnosis in Learning Disabilities
    Role: Principal Investigator
    R01HD022223     (OLSON, RICHARD K)Jan 1, 1987 - Dec 31, 1999
    NIH
    COMPUTER-SPEECH FEEDBACK IN TEXT FOR DYSLEXIC CHILDREN
    Role: Principal Investigator

    Collapse Bibliographic 
    Collapse selected publications
    Publications listed below are automatically derived from MEDLINE/PubMed and other sources, which might result in incorrect or missing publications. Faculty can login to make corrections and additions.
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    1. Doust C, Fontanillas P, Eising E, Gordon SD, Wang Z, Alag?z G, Molz B, Pourcain BS, Francks C, Marioni RE, Zhao J, Paracchini S, Talcott JB, Monaco AP, Stein JF, Gruen JR, Olson RK, Willcutt EG, DeFries JC, Pennington BF, Smith SD, Wright MJ, Martin NG, Auton A, Bates TC, Fisher SE, Luciano M. Author Correction: Discovery of 42 genome-wide significant loci associated with dyslexia. Nat Genet. 2023 Mar; 55(3):520. PMID: 36823321.
      View in: PubMed
    2. Doust C, Fontanillas P, Eising E, Gordon SD, Wang Z, Alag?z G, Molz B, Pourcain BS, Francks C, Marioni RE, Zhao J, Paracchini S, Talcott JB, Monaco AP, Stein JF, Gruen JR, Olson RK, Willcutt EG, DeFries JC, Pennington BF, Smith SD, Wright MJ, Martin NG, Auton A, Bates TC, Fisher SE, Luciano M. Discovery of 42 genome-wide significant loci associated with dyslexia. Nat Genet. 2022 11; 54(11):1621-1629. PMID: 36266505.
      View in: PubMed
    3. Eising E, Mirza-Schreiber N, de Zeeuw EL, Wang CA, Truong DT, Allegrini AG, Shapland CY, Zhu G, Wigg KG, Gerritse ML, Molz B, Alag?z G, Gialluisi A, Abbondanza F, Rimfeld K, van Donkelaar M, Liao Z, Jansen PR, Andlauer TFM, Bates TC, Bernard M, Blokland K, Bonte M, B?rglum AD, Bourgeron T, Brandeis D, Ceroni F, Cs?pe V, Dale PS, de Jong PF, DeFries JC, D?monet JF, Demontis D, Feng Y, Gordon SD, Guger SL, Hayiou-Thomas ME, Hern?ndez-Cabrera JA, Hottenga JJ, Hulme C, Kere J, Kerr EN, Koomar T, Landerl K, Leonard GT, Lovett MW, Lyytinen H, Martin NG, Martinelli A, Maurer U, Michaelson JJ, Moll K, Monaco AP, Morgan AT, N?then MM, Pausova Z, Pennell CE, Pennington BF, Price KM, Rajagopal VM, Ramus F, Richer L, Simpson NH, Smith SD, Snowling MJ, Stein J, Strug LJ, Talcott JB, Tiemeier H, van der Schroeff MP, Verhoef E, Watkins KE, Wilkinson M, Wright MJ, Barr CL, Boomsma DI, Carreiras M, Franken MJ, Gruen JR, Luciano M, M?ller-Myhsok B, Newbury DF, Olson RK, Paracchini S, Paus T, Plomin R, Reilly S, Schulte-K?rne G, Tomblin JB, van Bergen E, Whitehouse AJO, Willcutt EG, St Pourcain B, Francks C, Fisher SE. Genome-wide analyses of individual differences in quantitatively assessed reading- and language-related skills in up to 34,000 people. Proc Natl Acad Sci U S A. 2022 08 30; 119(35):e2202764119. PMID: 35998220.
      View in: PubMed
    4. Wakeman HN, Wadsworth SJ, Olson RK, DeFries JC, Pennington BF, Willcutt EG. Mathematics Difficulties and Psychopathology in School-Age Children. J Learn Disabil. 2023 Mar-Apr; 56(2):116-131. PMID: 35466804.
      View in: PubMed
    5. Wakeman HN, Leopold DR, Olson RK, Willcutt EG. Modeling the Speeded Determinants of Adolescents' Academic and Attentional Functioning. Dev Neuropsychol. 2022 Mar-Apr; 47(2):61-77. PMID: 34894917.
      View in: PubMed
    6. Peterson RL, McGrath LM, Willcutt EG, Keenan JM, Olson RK, Pennington BF. How Specific Are Learning Disabilities? J Learn Disabil. 2021 11; 54(6):466-483. PMID: 33446025.
      View in: PubMed
    7. Gialluisi A, Andlauer TFM, Mirza-Schreiber N, Moll K, Becker J, Hoffmann P, Ludwig KU, Czamara D, Pourcain BS, Honbolyg? F, T?th D, Cs?pe V, Huguet G, Chaix Y, Iannuzzi S, Demonet JF, Morris AP, Hulslander J, Willcutt EG, DeFries JC, Olson RK, Smith SD, Pennington BF, Vaessen A, Maurer U, Lyytinen H, Peyrard-Janvid M, Lepp?nen PHT, Brandeis D, Bonte M, Stein JF, Talcott JB, Fauchereau F, Wilcke A, Kirsten H, M?ller B, Francks C, Bourgeron T, Monaco AP, Ramus F, Landerl K, Kere J, Scerri TS, Paracchini S, Fisher SE, Schumacher J, N?then MM, M?ller-Myhsok B, Schulte-K?rne G. Genome-wide association study reveals new insights into the heritability and genetic correlates of developmental dyslexia. Mol Psychiatry. 2021 07; 26(7):3004-3017. PMID: 33057169.
      View in: PubMed
    8. Slomowitz RF, Narayan AJ, Pennington BF, Olson RK, DeFries JC, Willcutt EG, McGrath LM. In Search of Cognitive Promotive and Protective Factors for Word Reading. Sci Stud Read. 2021; 25(5):397-416. PMID: 34650325.
      View in: PubMed
    9. Little CW, Larsen S, Byrne B, Logan JAR, Olson RK, Coventry WL. Exploring the Influence of Early Childhood Education and Care on the Etiology of Achievement. Behav Genet. 2020 11; 50(6):387-400. PMID: 32797343.
      View in: PubMed
    10. Compton DL, Gilbert JK, Kearns DM, Olson RK. Using an item-specific predictor to test the dimensionality of the orthographic choice task. Ann Dyslexia. 2020 07; 70(2):243-258. PMID: 32712817.
      View in: PubMed
    11. Byrne B, Little CW, Olson RK, Larsen SA, Coventry WL, Weymouth R. Comment on Asbury and Wai (2019), "Viewing education policy through a genetic lens," Journal of School Choice. J Sch Choice. 2020; 14(3):501-515. PMID: 33727903.
      View in: PubMed
    12. Larsen SA, Little CW, Grasby K, Byrne B, Olson RK, Coventry WL. The Academic Development Study of Australian Twins (ADSAT): Research Aims and Design. Twin Res Hum Genet. 2020 06; 23(3):165-173. PMID: 32482186.
      View in: PubMed
    13. McGowan D, Little CW, Coventry WL, Corley R, Olson RK, Samuelsson S, Byrne B. Differential Influences of Genes and Environment Across the Distribution of Reading Ability. Behav Genet. 2019 09; 49(5):425-431. PMID: 31385189.
      View in: PubMed
    14. MacDonald B, Pennington BF, Willcutt EG, Dmitrieva J, Samuelsson S, Byrne B, Olson RK. Cross-Country Differences in Parental Reporting of Symptoms of ADHD. J Cross Cult Psychol. 2019 Jul; 50(6):806-824. PMID: 31303678.
      View in: PubMed
    15. Willcutt EG, McGrath LM, Pennington BF, Keenan JM, DeFries JC, Olson RK, Wadsworth SJ. Understanding Comorbidity Between Specific Learning Disabilities. New Dir Child Adolesc Dev. 2019 May; 2019(165):91-109. PMID: 31070302.
      View in: PubMed
    16. Truong DT, Adams AK, Paniagua S, Frijters JC, Boada R, Hill DE, Lovett MW, Mahone EM, Willcutt EG, Wolf M, Defries JC, Gialluisi A, Francks C, Fisher SE, Olson RK, Pennington BF, Smith SD, Bosson-Heenan J, Gruen JR. Multivariate genome-wide association study of rapid automatised naming and rapid alternating stimulus in Hispanic American and African-American youth. J Med Genet. 2019 08; 56(8):557-566. PMID: 30995994.
      View in: PubMed
    17. Gialluisi A, Andlauer TFM, Mirza-Schreiber N, Moll K, Becker J, Hoffmann P, Ludwig KU, Czamara D, St Pourcain B, Brandler W, Honbolyg? F, T?th D, Cs?pe V, Huguet G, Morris AP, Hulslander J, Willcutt EG, DeFries JC, Olson RK, Smith SD, Pennington BF, Vaessen A, Maurer U, Lyytinen H, Peyrard-Janvid M, Lepp?nen PHT, Brandeis D, Bonte M, Stein JF, Talcott JB, Fauchereau F, Wilcke A, Francks C, Bourgeron T, Monaco AP, Ramus F, Landerl K, Kere J, Scerri TS, Paracchini S, Fisher SE, Schumacher J, N?then MM, M?ller-Myhsok B, Schulte-K?rne G. Genome-wide association scan identifies new variants associated with a cognitive predictor of dyslexia. Transl Psychiatry. 2019 02 11; 9(1):77. PMID: 30741946.
      View in: PubMed
    18. Leopold DR, Christopher ME, Olson RK, Petrill SA, Willcutt EG. Invariance of ADHD Symptoms Across Sex and Age: a Latent Analysis of ADHD and Impairment Ratings from Early Childhood into Adolescence. J Abnorm Child Psychol. 2019 01; 47(1):21-34. PMID: 29691720.
      View in: PubMed
    19. Becker SP, Burns GL, Leopold DR, Olson RK, Willcutt EG. Differential impact of trait sluggish cognitive tempo and ADHD inattention in early childhood on adolescent functioning. J Child Psychol Psychiatry. 2018 10; 59(10):1094-1104. PMID: 29957822.
      View in: PubMed
    20. Gould KL, Coventry WL, Olson RK, Byrne B. Gene-Environment Interactions in ADHD: The Roles of SES and Chaos. J Abnorm Child Psychol. 2018 02; 46(2):251-263. PMID: 28283857.
      View in: PubMed
    21. Lukowski SL, Rosenberg-Lee M, Thompson LA, Hart SA, Willcutt EG, Olson RK, Petrill SA, Pennington BF. Approximate Number Sense Shares Etiological Overlap with Mathematics and General Cognitive Ability. Intelligence. 2017 Nov; 65:67-74. PMID: 29867283.
      View in: PubMed
    22. Adams AK, Smith SD, Truong DT, Willcutt EG, Olson RK, DeFries JC, Pennington BF, Gruen JR. Enrichment of putatively damaging rare variants in the DYX2 locus and the reading-related genes CCDC136 and FLNC. Hum Genet. 2017 11; 136(11-12):1395-1405. PMID: 28866788.
      View in: PubMed
    23. Grasby KL, Coventry WL, Byrne B, Olson RK. Little Evidence That Socioeconomic Status Modifies Heritability of Literacy and Numeracy in Australia. Child Dev. 2019 03; 90(2):623-637. PMID: 28832969.
      View in: PubMed
    24. Peterson RL, Arnett AB, Pennington BF, Byrne B, Samuelsson S, Olson RK. Literacy acquisition influences children's rapid automatized naming. Dev Sci. 2018 05; 21(3):e12589. PMID: 28812316.
      View in: PubMed
    25. Burrows T, Goldman S, Olson RK, Byrne B, Coventry WL. Associations between selected dietary behaviours and academic achievement: A study of Australian school aged children. Appetite. 2017 09 01; 116:372-380. PMID: 28487248.
      View in: PubMed
    26. Arnett AB, Pennington BF, Peterson RL, Willcutt EG, DeFries JC, Olson RK. Explaining the sex difference in dyslexia. J Child Psychol Psychiatry. 2017 Jun; 58(6):719-727. PMID: 28176347.
      View in: PubMed
    27. Livingstone LT, Coventry WL, Corley RP, Willcutt EG, Samuelsson S, Olson RK, Byrne B. Does the Environment Have an Enduring Effect on ADHD? A Longitudinal Study of Monozygotic Twin Differences in Children. J Abnorm Child Psychol. 2016 11; 44(8):1487-1501. PMID: 26993487.
      View in: PubMed
    28. Wadsworth SJ, DeFries JC, Willcutt EG, Pennington BF, Olson RK. Genetic Etiologies of Comorbidity and Stability for Reading Difficulties and ADHD: A Replication Study. Twin Res Hum Genet. 2016 12; 19(6):647-651. PMID: 27748217.
      View in: PubMed
    29. Treiman R, Kessler B, Pollo TC, Byrne B, Olson RK. Measures of Kindergarten Spelling and Their Relations to Later Spelling Performance. Sci Stud Read. 2016; 20(5):349-362. PMID: 27761101.
      View in: PubMed
    30. Grasby KL, Coventry WL, Byrne B, Olson RK, Medland SE. Genetic and Environmental Influences on Literacy and Numeracy Performance in Australian School Children in Grades 3, 5, 7, and 9. Behav Genet. 2016 Sep; 46(5):627-648. PMID: 27276978.
      View in: PubMed
    31. Gialluisi A, Visconti A, Willcutt EG, Smith SD, Pennington BF, Falchi M, DeFries JC, Olson RK, Francks C, Fisher SE. Investigating the effects of copy number variants on reading and language performance. J Neurodev Disord. 2016; 8:17. PMID: 27186239.
      View in: PubMed
    32. Christopher ME, Keenan JM, Hulslander J, DeFries JC, Miyake A, Wadsworth SJ, Willcutt E, Pennington B, Olson RK. The genetic and environmental etiologies of the relations between cognitive skills and components of reading ability. J Exp Psychol Gen. 2016 Apr; 145(4):451-66. PMID: 26974208.
      View in: PubMed
    33. Peterson RL, Boada R, McGrath LM, Willcutt EG, Olson RK, Pennington BF. Cognitive Prediction of Reading, Math, and Attention: Shared and Unique Influences. J Learn Disabil. 2017 Jul/Aug; 50(4):408-421. PMID: 26825667.
      View in: PubMed
    34. Leopold DR, Christopher ME, Burns GL, Becker SP, Olson RK, Willcutt EG. Attention-deficit/hyperactivity disorder and sluggish cognitive tempo throughout childhood: temporal invariance and stability from preschool through ninth grade. J Child Psychol Psychiatry. 2016 09; 57(9):1066-74. PMID: 26749438.
      View in: PubMed
    35. Powers NR, Eicher JD, Miller LL, Kong Y, Smith SD, Pennington BF, Willcutt EG, Olson RK, Ring SM, Gruen JR. The regulatory element READ1 epistatically influences reading and language, with both deleterious and protective alleles. J Med Genet. 2016 Mar; 53(3):163-71. PMID: 26660103.
      View in: PubMed
    36. Wadsworth SJ, DeFries JC, Willcutt EG, Pennington BF, Olson RK. The Colorado Longitudinal Twin Study of Reading Difficulties and ADHD: Etiologies of Comorbidity and Stability. Twin Res Hum Genet. 2015 Dec; 18(6):755-61. PMID: 26537134.
      View in: PubMed
    37. Grasby KL, Byrne B, Olson RK. Validity of large-scale reading tests: A phenotypic and behaviour-genetic analysis. Aust J Educ. 2015 Apr; 59(1):5-21. PMID: 27721516.
      View in: PubMed
    38. Soden B, Christopher ME, Hulslander J, Olson RK, Cutting L, Keenan JM, Thompson LA, Wadsworth SJ, Willcutt EG, Petrill SA. Longitudinal stability in reading comprehension is largely heritable from grades 1 to 6. PLoS One. 2015; 10(1):e0113807. PMID: 25602760.
      View in: PubMed
    39. Elw?r ?, Gustafson S, Byrne B, Olson RK, Keenan JM, Samuelsson S. A retrospective longitudinal study of cognitive and language skills in poor reading comprehension. Scand J Psychol. 2015 Apr; 56(2):157-66. PMID: 25581078.
      View in: PubMed
    40. Olson RK, Keenan JM, Byrne B, Samuelsson S. Why do Children Differ in Their Development of Reading and Related Skills? Sci Stud Read. 2014 Dec 31; 18(1):38-54. PMID: 25104901.
      View in: PubMed
    41. Keenan JM, Hua AN, Meenan CE, Pennington BF, Willcutt E, Olson RK. Issues in Identifying Poor Comprehenders. Annee Psychol. 2014 Dec 01; 114(4):753-777. PMID: 25937640.
      View in: PubMed
    42. Arnett AB, Pennington BF, Willcutt EG, DeFries JC, Olson RK. Sex differences in ADHD symptom severity. J Child Psychol Psychiatry. 2015 Jun; 56(6):632-9. PMID: 25283790.
      View in: PubMed
    43. Christopher ME, Hulslander J, Byrne B, Samuelsson S, Keenan JM, Pennington B, DeFries JC, Wadsworth SJ, Willcutt E, Olson RK. Genetic and environmental etiologies of the longitudinal relations between prereading skills and reading. Child Dev. 2015 Mar-Apr; 86(2):342-61. PMID: 25263167.
      View in: PubMed
    44. Eicher JD, Powers NR, Miller LL, Mueller KL, Mascheretti S, Marino C, Willcutt EG, DeFries JC, Olson RK, Smith SD, Pennington BF, Tomblin JB, Ring SM, Gruen JR. Characterization of the DYX2 locus on chromosome 6p22 with reading disability, language impairment, and IQ. Hum Genet. 2014 Jul; 133(7):869-81. PMID: 24509779.
      View in: PubMed
    45. Willcutt EG, Chhabildas N, Kinnear M, DeFries JC, Olson RK, Leopold DR, Keenan JM, Pennington BF. The internal and external validity of sluggish cognitive tempo and its relation with DSM-IV ADHD. J Abnorm Child Psychol. 2014 Jan; 42(1):21-35. PMID: 24122408.
      View in: PubMed
    46. Peterson RL, Pennington BF, Olson RK, Wadsworth S. Longitudinal Stability of Phonological and Surface Subtypes of Developmental Dyslexia. Sci Stud Read. 2014; 18(5):347-362. PMID: 25429194.
      View in: PubMed
    47. Christopher ME, Hulslander J, Byrne B, Samuelsson S, Keenan JM, Pennington B, Defries JC, Wadsworth SJ, Willcutt E, Olson RK. The genetic and environmental etiologies of individual differences in early reading growth in Australia, the United States, and Scandinavia. J Exp Child Psychol. 2013 Jul; 115(3):453-67. PMID: 23665180.
      View in: PubMed
    48. Miller AC, Keenan JM, Betjemann RS, Willcutt EG, Pennington BF, Olson RK. Reading comprehension in children with ADHD: cognitive underpinnings of the centrality deficit. J Abnorm Child Psychol. 2013 Apr; 41(3):473-83. PMID: 23054132.
      View in: PubMed
    49. Fujisawa KK, Wadsworth SJ, Kakihana S, Olson RK, Defries JC, Byrne B, Ando J. A multivariate twin study of early literacy in Japanese Kana. Learn Individ Differ. 2013 Apr 01; 24:160-167. PMID: 23997545.
      View in: PubMed
    50. Elw?r S, Keenan JM, Olson RK, Byrne B, Samuelsson S. Longitudinal stability and predictors of poor oral comprehenders and poor decoders. J Exp Child Psychol. 2013 Jul; 115(3):497-516. PMID: 23528975.
      View in: PubMed
    51. Willcutt EG, Petrill SA, Wu S, Boada R, Defries JC, Olson RK, Pennington BF. Comorbidity between reading disability and math disability: concurrent psychopathology, functional impairment, and neuropsychological functioning. J Learn Disabil. 2013 Nov-Dec; 46(6):500-16. PMID: 23449727.
      View in: PubMed
    52. Byrne B, Wadsworth SJ, Boehme K, Talk AC, Coventry WL, Olson RK, Samuelsson S, Corley R. Multivariate genetic analysis of learning and early reading development. Sci Stud Read. 2013 Jan 01; 17(3):224-242. PMID: 23626456.
      View in: PubMed
    53. Christopher ME, Hulslander J, Byrne B, Samuelsson S, Keenan JM, Pennington B, Defries JC, Wadsworth SJ, Willcutt E, Olson RK. Modeling the Etiology of Individual Differences in Early Reading Development: Evidence for Strong Genetic Influences. Sci Stud Read. 2013; 17(5):350-368. PMID: 24489459.
      View in: PubMed
    54. Peterson RL, Pennington BF, Samuelsson S, Byrne B, Olson RK. Shared etiology of phonological memory and vocabulary deficits in school-age children. J Speech Lang Hear Res. 2013 Aug; 56(4):1249-59. PMID: 23275423.
      View in: PubMed
    55. Arnett AB, Pennington BF, Willcutt E, Dmitrieva J, Byrne B, Samuelsson S, Olson RK. A cross-lagged model of the development of ADHD inattention symptoms and rapid naming speed. J Abnorm Child Psychol. 2012 Nov; 40(8):1313-26. PMID: 22581405.
      View in: PubMed
    56. Peterson RL, Pennington BF, Olson RK. Subtypes of developmental dyslexia: testing the predictions of the dual-route and connectionist frameworks. Cognition. 2013 Jan; 126(1):20-38. PMID: 23010562.
      View in: PubMed
    57. Anthoni H, Sucheston LE, Lewis BA, Tapia-P?ez I, Fan X, Zucchelli M, Taipale M, Stein CM, Hokkanen ME, Castr?n E, Pennington BF, Smith SD, Olson RK, Tomblin JB, Schulte-K?rne G, N?then M, Schumacher J, M?ller-Myhsok B, Hoffmann P, Gilger JW, Hynd GW, Nopola-Hemmi J, Leppanen PH, Lyytinen H, Schoumans J, Nordenskj?ld M, Spencer J, Stanic D, Boon WC, Simpson E, M?kel? S, Gustafsson J?, Peyrard-Janvid M, Iyengar S, Kere J. The aromatase gene CYP19A1: several genetic and functional lines of evidence supporting a role in reading, speech and language. Behav Genet. 2012 Jul; 42(4):509-27. PMID: 22426781.
      View in: PubMed
    58. Christopher ME, Miyake A, Keenan JM, Pennington B, DeFries JC, Wadsworth SJ, Willcutt E, Olson RK. Predicting word reading and comprehension with executive function and speed measures across development: a latent variable analysis. J Exp Psychol Gen. 2012 Aug; 141(3):470-488. PMID: 22352396.
      View in: PubMed
    59. Astrom RL, Wadsworth SJ, Olson RK, Willcutt EG, Defries JC. Genetic and environmental etiologies of reading difficulties: DeFries-Fulker analysis of reading performance data from twin pairs and their nontwin siblings. Learn Individ Differ. 2012 Jun 01; 22(3):365-369. PMID: 22927712.
      View in: PubMed
    60. Wadsworth SJ, Olson RK, Willcutt EG, DeFries JC. Multiple regression analysis of reading performance data from twin pairs with reading difficulties and nontwin siblings: the augmented model. Twin Res Hum Genet. 2012 Feb; 15(1):116-9. PMID: 22784461.
      View in: PubMed
    61. Arnett AB, Pennington BF, Friend A, Willcutt EG, Byrne B, Samuelsson S, Olson RK. The SWAN captures variance at the negative and positive ends of the ADHD symptom dimension. J Atten Disord. 2013 Feb; 17(2):152-62. PMID: 22173148.
      View in: PubMed
    62. Pennington BF, Santerre-Lemmon L, Rosenberg J, MacDonald B, Boada R, Friend A, Leopold DR, Samuelsson S, Byrne B, Willcutt EG, Olson RK. Individual prediction of dyslexia by single versus multiple deficit models. J Abnorm Psychol. 2012 Feb; 121(1):212-24. PMID: 22022952.
      View in: PubMed
    63. Rosenberg J, Pennington BF, Willcutt EG, Olson RK. Gene by environment interactions influencing reading disability and the inattentive symptom dimension of attention deficit/hyperactivity disorder. J Child Psychol Psychiatry. 2012 Mar; 53(3):243-51. PMID: 21884522.
      View in: PubMed
    64. Olson RK, Hulslander J, Christopher M, Keenan JM, Wadsworth SJ, Willcutt EG, Pennington BF, DeFries JC. Genetic and environmental influences on writing and their relations to language and reading. Ann Dyslexia. 2013 Apr; 63(1):25-43. PMID: 21842316.
      View in: PubMed
    65. Coventry WL, Byrne B, Olson RK, Corley R, Samuelsson S. Dynamic and static assessment of phonological awareness in preschool: a behavior-genetic study. J Learn Disabil. 2011 Jul-Aug; 44(4):322-9. PMID: 21571699.
      View in: PubMed
    66. Astrom RL, Wadsworth SJ, Olson RK, Willcutt EG, DeFries JC. DeFries-Fulker analysis of longitudinal reading performance data from twin pairs ascertained for reading difficulties and from their nontwin siblings. Behav Genet. 2011 Sep; 41(5):660-7. PMID: 21259040.
      View in: PubMed
    67. Bidwell LC, Willcutt EG, McQueen MB, DeFries JC, Olson RK, Smith SD, Pennington BF. A family based association study of DRD4, DAT1, and 5HTT and continuous traits of attention-deficit hyperactivity disorder. Behav Genet. 2011 Jan; 41(1):165-74. PMID: 21207241.
      View in: PubMed
    68. Olson RK, Keenan JM, Byrne B, Samuelsson S, Coventry WL, Corley R, Wadsworth SJ, Willcutt EG, Defries JC, Pennington BF, Hulslander J. Genetic and Environmental Influences on Vocabulary and Reading Development. Sci Stud Read. 2011; 15(1):26-46. PMID: 21132077.
      View in: PubMed
    69. Betjemann RS, Keenan JM, Olson RK, Defries JC. Choice of Reading Comprehension Test Influences the Outcomes of Genetic Analyses. Sci Stud Read. 2011 Jan 01; 15(4):363-382. PMID: 21804757.
      View in: PubMed
    70. McGrath LM, Pennington BF, Shanahan MA, Santerre-Lemmon LE, Barnard HD, Willcutt EG, Defries JC, Olson RK. A multiple deficit model of reading disability and attention-deficit/hyperactivity disorder: searching for shared cognitive deficits. J Child Psychol Psychiatry. 2011 May; 52(5):547-57. PMID: 21126246.
      View in: PubMed
    71. Scerri TS, Paracchini S, Morris A, MacPhie IL, Talcott J, Stein J, Smith SD, Pennington BF, Olson RK, DeFries JC, Monaco AP, Richardson AJ. Identification of candidate genes for dyslexia susceptibility on chromosome 18. PLoS One. 2010 Oct 28; 5(10):e13712. PMID: 21060895.
      View in: PubMed
    72. Smith SD, Grigorenko E, Willcutt E, Pennington BF, Olson RK, DeFries JC. Etiologies and molecular mechanisms of communication disorders. J Dev Behav Pediatr. 2010 Sep; 31(7):555-63. PMID: 20814255.
      View in: PubMed
    73. Willcutt EG, Pennington BF, Duncan L, Smith SD, Keenan JM, Wadsworth S, Defries JC, Olson RK. Understanding the complex etiologies of developmental disorders: behavioral and molecular genetic approaches. J Dev Behav Pediatr. 2010 Sep; 31(7):533-44. PMID: 20814254.
      View in: PubMed
    74. Willcutt EG, Betjemann RS, McGrath LM, Chhabildas NA, Olson RK, DeFries JC, Pennington BF. Etiology and neuropsychology of comorbidity between RD and ADHD: the case for multiple-deficit models. Cortex. 2010 Nov-Dec; 46(10):1345-61. PMID: 20828676.
      View in: PubMed
    75. Ebejer JL, Coventry WL, Byrne B, Willcutt EG, Olson RK, Corley R, Samuelsson S. Genetic and Environmental Influences on Inattention, Hyperactivity-Impulsivity, and Reading: Kindergarten to Grade 2. Sci Stud Read. 2010 Jul; 14(4):293-316. PMID: 20823940.
      View in: PubMed
    76. Hulslander J, Olson RK, Willcutt EG, Wadsworth SJ. Longitudinal Stability of Reading-Related Skills and their Prediction of Reading Development. Sci Stud Read. 2010 Mar 01; 14(2):111-136. PMID: 20563241.
      View in: PubMed
    77. Friend A, Olson RK. Phonological Spelling and Reading Deficits in Children with Spelling Disabilities. Sci Stud Read. 2010 Jan; 12(1):90-105. PMID: 20585591.
      View in: PubMed
    78. Coventry WL, Byrne B, Coleman M, Olson RK, Corley R, Willcutt E, Samuelsson S. Does classroom separation affect twins' reading ability in the early years of school? Twin Res Hum Genet. 2009 Oct; 12(5):455-61. PMID: 19803773.
      View in: PubMed
    79. Hawke JL, Olson RK, Willcut EG, Wadsworth SJ, DeFries JC. Gender ratios for reading difficulties. Dyslexia. 2009 Aug; 15(3):239-42. PMID: 19367616.
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    81. Friend A, DeFries JC, Olson RK, Pennington B, Harlaar N, Byrne B, Samuelsson S, Willcutt EG, Wadsworth SJ, Corley R, Keenan JM. Heritability of high reading ability and its interaction with parental education. Behav Genet. 2009 Jul; 39(4):427-36. PMID: 19296213.
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    83. Friend A, DeFries JC, Olson RK. Parental education moderates genetic influences on reading disability. Psychol Sci. 2008 Nov; 19(11):1124-30. PMID: 19076484.
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    84. Samuelsson S, Byrne B, Olson RK, Hulslander J, Wadsworth S, Corley R, Willcutt EG, Defries JC. Response to early literacy instruction in the United States, Australia, and Scandinavia: A behavioral-genetic analysis. Learn Individ Differ. 2008; 18(3):289-295. PMID: 19122888.
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    85. Wadsworth SJ, DeFries JC, Olson RK, Willcutt EG. Colorado longitudinal twin study of reading disability. Ann Dyslexia. 2007 Dec; 57(2):139-60. PMID: 18060583.
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    86. Willcutt EG, Pennington BF, Olson RK, DeFries JC. Understanding comorbidity: a twin study of reading disability and attention-deficit/hyperactivity disorder. Am J Med Genet B Neuropsychiatr Genet. 2007 Sep 05; 144B(6):709-14. PMID: 17440942.
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    87. Phinney E, Pennington BF, Olson R, Filley CM, Filipek PA. Brain structure correlates of component reading processes: implications for reading disability. Cortex. 2007 Aug; 43(6):777-91. PMID: 17710829.
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    88. Friend A, DeFries JC, Wadsworth SJ, Olson RK. Genetic and environmental influences on word recognition and spelling deficits as a function of age. Behav Genet. 2007 May; 37(3):477-86. PMID: 17345157.
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    89. Olson RK. Genes, environment, and dyslexia. The 2005 Norman Geschwind Memorial Lecture. Ann Dyslexia. 2006 Dec; 56(2):205-38. PMID: 17849199.
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    90. Shanahan MA, Pennington BF, Yerys BE, Scott A, Boada R, Willcutt EG, Olson RK, DeFries JC. Processing speed deficits in attention deficit/hyperactivity disorder and reading disability. J Abnorm Child Psychol. 2006 Oct; 34(5):585-602. PMID: 16850284.
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    91. Meng H, Smith SD, Hager K, Held M, Liu J, Olson RK, Pennington BF, DeFries JC, Gelernter J, O'Reilly-Pol T, Somlo S, Skudlarski P, Shaywitz SE, Shaywitz BA, Marchione K, Wang Y, Paramasivam M, LoTurco JJ, Page GP, Gruen JR. DCDC2 is associated with reading disability and modulates neuronal development in the brain. Proc Natl Acad Sci U S A. 2005 Nov 22; 102(47):17053-8. PMID: 16278297.
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    92. Meng H, Hager K, Held M, Page GP, Olson RK, Pennington BF, DeFries JC, Smith SD, Gruen JR. TDT-association analysis of EKN1 and dyslexia in a Colorado twin cohort. Hum Genet. 2005 Oct; 118(1):87-90. PMID: 16133186.
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    93. Willcutt EG, Pennington BF, Olson RK, Chhabildas N, Hulslander J. Neuropsychological analyses of comorbidity between reading disability and attention deficit hyperactivity disorder: in search of the common deficit. Dev Neuropsychol. 2005; 27(1):35-78. PMID: 15737942.
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    94. Francks C, Paracchini S, Smith SD, Richardson AJ, Scerri TS, Cardon LR, Marlow AJ, MacPhie IL, Walter J, Pennington BF, Fisher SE, Olson RK, DeFries JC, Stein JF, Monaco AP. A 77-kilobase region of chromosome 6p22.2 is associated with dyslexia in families from the United Kingdom and from the United States. Am J Hum Genet. 2004 Dec; 75(6):1046-58. PMID: 15514892.
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    95. Hulslander J, Talcott J, Witton C, DeFries J, Pennington B, Wadsworth S, Willcutt E, Olson R. Sensory processing, reading, IQ, and attention. J Exp Child Psychol. 2004 Jul; 88(3):274-95. PMID: 15203301.
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    96. Tiu RD, Wadsworth SJ, Olson RK, DeFries JC. Causal models of reading disability: a twin study. Twin Res. 2004 Jun; 7(3):275-83. PMID: 15193172.
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    97. Deffenbacher KE, Kenyon JB, Hoover DM, Olson RK, Pennington BF, DeFries JC, Smith SD. Refinement of the 6p21.3 quantitative trait locus influencing dyslexia: linkage and association analyses. Hum Genet. 2004 Jul; 115(2):128-38. PMID: 15138886.
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    98. Gay?n J, Olson RK. Genetic and environmental influences on individual differences in printed word recognition. J Exp Child Psychol. 2003 Feb; 84(2):97-123. PMID: 12609495.
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    99. Olson RK. Dyslexia: nature and nurture. Dyslexia. 2002 Jul-Sep; 8(3):143-59. PMID: 12222731.
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    100. Knopik VS, Smith SD, Cardon L, Pennington B, Gayan J, Olson RK, DeFries JC. Differential genetic etiology of reading component processes as a function of IQ. Behav Genet. 2002 May; 32(3):181-98. PMID: 12141780.
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    101. Willcutt EG, Pennington BF, Smith SD, Cardon LR, Gay?n J, Knopik VS, Olson RK, DeFries JC. Quantitative trait locus for reading disability on chromosome 6p is pleiotropic for attention-deficit/hyperactivity disorder. Am J Med Genet. 2002 Apr 08; 114(3):260-8. PMID: 11920845.
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    102. Francks C, Fisher SE, Olson RK, Pennington BF, Smith SD, DeFries JC, Monaco AP. Fine mapping of the chromosome 2p12-16 dyslexia susceptibility locus: quantitative association analysis and positional candidate genes SEMA4F and OTX1. Psychiatr Genet. 2002 Mar; 12(1):35-41. PMID: 11901358.
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    103. Fisher SE, Francks C, Marlow AJ, MacPhie IL, Newbury DF, Cardon LR, Ishikawa-Brush Y, Richardson AJ, Talcott JB, Gay?n J, Olson RK, Pennington BF, Smith SD, DeFries JC, Stein JF, Monaco AP. Independent genome-wide scans identify a chromosome 18 quantitative-trait locus influencing dyslexia. Nat Genet. 2002 Jan; 30(1):86-91. PMID: 11743577.
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